TYPE: Costa Rica. San José: Pérez Zeledón, Páramo, San Ramón Sur, toward Río Berlin, ca. 9º25'N, 83º44'W, 1420-1640 m, collected by Esteban Víquez J., flowered in cultivation in the collection of Marta Herra, 29 May 2004, F. Pupulin 5236 (holotype, USJ; isotype, CR, USJ-spirit; illustration of type; photo of the flowers from the plant that served as the holotype).