TYPE: Nicaragua. Matagalpa: Cordillera Central, area near "Jinotega Rock Quarry", 5 km north of Santa María Ostuma, 1500 m, 8 Jan 1963, L.O. Williams, A. Molina R. & T.P. Williams as A.H. Heller 23373 (holotype, SEL, (photo, AMO); illustration of type).