TYPE: Panamá. Darién: Parque Nacional del Darién Ridge between N & S branches of Río Púcuro, across river from old Kuna village of Tacarcuna; ca. 18 km E of Púcuro, 8°04' N 7r16' W, 600-1000 m, 21 October 1987, B. Hammel, G. de Nevers, H. Cuadros & H. Herrera 16342 (holotype, MO; illustration of type).