TYPE: Nicaragua. Dept. Grenada: western slope of Volcán Mombacho, road and trail above Finca Santa Ana, reservoir somewhat above Plan de las Flores, ca. 11º49'-50'N, 85º58'W, elev. ca. 950-1150 m, 1 Oct 1977, W.D. Stevens & B.A. Krukoff 4358 (holotype, SEL; isotype, MO; illustration of type).