TYPE: Costa Rica. Cartago: in forest along creek debouching into Río Grande de Orosí from the east, ca. 1 km upstream from confluence of Quebrada Salta, Tapantí, 9º43'N 83º47'W, 1500-1700 m, 12 Jul 1984, Michael H. Grayum & Pam Sleeper 3446 (holotype, MO; illustration of type).