TYPE: Costa Rica. in den Wäldern von La Palma, 1700-2000 m, blühend im Juli 1895, A. Tonduz 9681 (holotype, B, destroyed; isotype, AMES; illustration of type, AMES; epitype designated by Atwood (1999), Costa Rica. La Palama, S. Ingram & K. Ferrell 770 (SEL).