TYPE: Costa Rica. Cartago: Turrialba, Moravia de Chirripó, Quebrada Tsipirí, 9º48'N 83º23'W, 1090 m, on steep slope along the Quebrada, lower montane rain forest, primary vegetation, 26 April 2002, F. Pupulin, H. Montealegre, M. Bonilla & J.C. Cervantes 3644 (holotype, USJ; illustration of type).